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First published 2008
ISBN-13 9780702028571
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 Preface
We believe that a good understanding of the physiol-
ogy of exercise is important for anyone involved in the
care, handling, training, feeding, breeding, or use of
horses. The care of horses, and athletic horses in par-
ticular, requires an understanding of the physiological
demands of exercise, which then provides a better
grasp of the requirements of competition and training.
The science of equine exercise physiology has pro-
gressed to the stage that it now provides a sound, sci-
entifi c basis for much of the nutrition, training, and
care of equine athletes. The current level of knowl-
edge, while still incomplete and imperfect, of the phys-
iological processes underlying the acute responses to
exercise and the mechanisms and effects of exercise
conditioning, provides a sound, fundamental under-
standing of the workings of the equine athlete. Con-
temporary equine exercise physiology comprises not
only the physiological responses to exercise and train-
ing, but also nutrition, biomechanics, behavior, and
pharmacology. This fundamental knowledge informs
our decisions regarding appropriate training, nutri-
tion, care, and treatment of the equine athlete.
The fi rst edition of
Equine Exercise Physiology
is a
response to requests by many readers of
Equine Sports
Medicine and Surgery: basic and clinical sciences of the
equine athlete
that we provide a text focused on equine
exercise physiology. This text is intended for anyone
involved with athletic horses, and in particular those
individuals who are not responsible for providing the
primary veterinary care of such horses. While the text
presupposes a good knowledge of physiology, we think
that it is written at a level that will allow most informed
individuals to gain much valuable information.
We thank the colleagues and students with whom
we have had the pleasure of working and who pro-
vided much of the knowledge contained within this
book. Our profound gratitude is extended to the
authors of sections of this book with an appreciation
of the effort that was required to compile anew com-
prehensive material on their designated topic. We
hope that they, and the readers, are pleased with the
fi nal product.
vii
Contributors
Dorothy M. Ainsworth
Professor
Dorothy Ainsworth
DVM PhD Dip ACVIM
Professor of Medicine
College of Veterinary Medicine
Cornell University
Ithaca, New York
USA
Raymond J. Geor
BVSc
MVSc PhD Dipl ACVIM
Professor and Paul Mellon Chair
Virginia Polytechnic and State University
Middleburg, Virginia
USA
Allen E. Goodship
BVSc PhD MRCVS
Professor of Orthopaedic Science
Royal Veterinary College
North Mymms, Hatfi eld
Director of Institute of Orthopaedics and
Musculoskeletal Science, University College London
UK
Eric Barrey
DVM PhD
INRA
Station de Génétique Quantitative et Appliquée
Jouy-en-Josas
France
Alicia L. Bertone
DVM PhD DipACVS
Professor, Trueman Family Endowed Chair
College of Veterinary Medicine
The Ohio State University
Columbus, Ohio
USA
Mary Elizabeth Gordon
PhD
Leader, New Product Development
Land O’Lakes Purina
Mills LLC
St Louis, Missouri
USA
Norm G. Ducharme
DVM MSc DipACVS
Professor of Surgery
College of Veterinary Medicine
Department of Clinical Sciences
Cornell University
Ithaca, New York
USA
Kenneth W. Hinchcliff
BVSc(Hons)
MS PhD Dipl
ACVIM
Professor and Dean
School of Veterinary Science
University of Melbourne
Melbourne
Australia
Howard H. Erickson
DVM PhD
Professor of Physiology
College of Veterinary Medicine
Kansas State University
Manhattan, Kansas
USA
Susan J. Holcombe
VMD
MS PhD DipACVS
ACVECC
Associate Professor
College of Veterinary Medicine
Department of Large Animal Clinical Sciences
Michigan State University
East Lansing, Michigan
USA
David Evans
BVSc PhD
Associate Professor
Faculty of Veterinary Science
University of Sydney
Sydney, New South Wales
Australia
David W. Horohov
PhD
William Robert Mills Chair in Equine Immunology
Department of Veterinary Science
University of Kentucky
Lexington, Kentucky
USA
viii
Contributors
ix
Seppo Hyyppä
DVM
Senior Scientist
Equine Research
Ypäjä, Finland
Richard J. Piercy
MA vetMB MS PhD DipACVIM MRCVS
Senior Lecturer in Equine Medicine
Royal Veterinary College
Hawkshead Lane
North Mymms
Hatfi eld
UK
Janene K. Kingston
BVSc
MVS DVSc PhD MACVSc
DipACVIM
Senior Lecturer in Equine Medicine
School of Veterinary Science
University of Queensland
Brisbane
Australia
David C. Poole
BSc MS PhD DSc FACSM
Professor of Kinesiology and Physiology
Department of Kinesiology and Anatomy and
Physiology
Kansas State University
131 Coles Hall
Manhattan, Kansas
USA
Michael I. Lindinger
PhD
Associate Professor
Department of Human Health
University of Guelph
Guelph, Ontario
Canada
A. Reeta Pösö
PhD
Professor of Veterinary Physiology
Department of Basic Veterinary Sciences
University of Helsinki
Finland
Mireia Lorenzo-Figueras
DVM PhD
Graduate Student
Veterinary Medical Teaching Hospital
University of California at Davis
Davis, California
USA
José Luis L. Rivero
DVM PhD
Professor of Veterinary Anatomy
Departmento de Anatomía
Universidad de Córdoba
Campus Rabanales-Sanidad Animal
Cordoba
Spain
L. Jill McCutcheon
BSc DVM PhD
Professor
Department of Biomedical Sciences and Pathology
Virginia–Maryland Regional College of Veterinary
Medicine
Blacksburg, Virginia
USA
Roger K.W. Smith
Professor of Equine Orthopaedics
Department of Veterinary Clinical Sciences
Royal Veterinary College
Hawkshead Lane
North Mymms
Hatfi eld
UK
Kenneth H. McKeever
PhD FACSM
Associate Professor and Associate Director
Department of Animal Science
Equine Science Center
Rutgers University
New Brunswick, New Jersey
USA
Amanda Waller
MSc PhD
Department of Human Biology and Nutritional
Sciences
University of Guelph
Guelph, Ontario
Canada
Alfred M. Merritt
DVM MS
Appleton Professor in Equine Studies
Large Animal Clinical Sciences
College of Veterinary Medicine
University of Florida
Gainesville, Florida
USA
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